1<!doctype html> 2<html lang="en"> 3 <head> 4 <meta charset="utf-8" /> 5 <link rel="icon" href="./favicon.svg" /> 6 <meta name="viewport" content="width=device-width, initial-scale=1" /> 7 8 <!-- Font preloading for better performance --> 9 <link rel="preconnect" href="https://fonts.googleapis.com" /> 10 <link rel="preconnect" href="https://fonts.gstatic.com" crossorigin /> 11 <link href="https://fonts.googleapis.com/css2?family=Lexend:wght@300;400;500;600;700&family=Lexend+Exa:wght@400&display=swap" rel="stylesheet" /> 12 13 <!-- Security headers --> 14 <meta http-equiv="X-Content-Type-Options" content="nosniff" /> 15 <meta http-equiv="X-XSS-Protection" content="1; mode=block" /> 16 <meta http-equiv="Referrer-Policy" content="strict-origin-when-cross-origin" /> 17 18 <meta name="robots" content="max-image-preview:large" /> 19 20
20<script src="https://mcp.figma.com/mcp/html-to-design/capture.js" async></script>
20 21 22 <!--dhnoa8--><meta name="description" content="Comprehensive list of scientific papers and research publications related to Sea Hero Quest, the spatial navigation game for Alzheimer's research."/> <meta property="og:type" content="website"/> <meta property="og:url" content="https://seaheroquest.com/papers"/> <meta property="og:title" content="Scientific Papers - Sea Hero Quest"/> <meta property="og:description" content="Comprehensive list of scientific papers and research publications related to Sea Hero Quest, the spatial navigation game for Alzheimer's research."/> <meta property="og:image" content="https://seaheroquest.com/shq-papers.png"/> <meta property="og:image:width" content="1200"/> <meta property="og:image:height" content="630"/> <meta property="og:image:alt" content="Sea Hero Quest scientific research papers and publications"/> <meta property="og:site_name" content="Sea Hero Quest"/> <meta property="og:locale" content="en_US"/> <meta property="twitter:card" content="summary_large_image"/> <meta property="twitter:url" content="https://seaheroquest.com/papers"/> <meta property="twitter:title" content="Scientific Papers - Sea Hero Quest"/> <meta property="twitter:description" content="Comprehensive list of scientific papers and research publications related to Sea Hero Quest, the spatial navigation game for Alzheimer's research."/> <meta property="twitter:image" content="https://seaheroquest.com/shq-papers.png"/> <meta property="twitter:image:alt" content="Sea Hero Quest scientific research papers and publications"/> <meta property="twitter:site" content="@seaheroquest"/> <meta property="twitter:creator" content="@seaheroquest"/> <meta name="keywords" content="Sea Hero Quest research, scientific papers, Alzheimer's research, spatial navigation studies, cognitive neuroscience, dementia research"/> <meta name="author" content="GLITCHERS LTD"/> <link rel="canonical" href="https://seaheroquest.com/papers"/>
22<script type="application/ld+json"> 23 { 24 "@context": "https://schema.org", 25 "@type": "CollectionPage", 26 "url": "https://seaheroquest.com/papers", 27 "name": "Sea Hero Quest â Scientific Papers", 28 "description": "Comprehensive list of peer-reviewed scientific papers and research publications from the Sea Hero Quest study on spatial navigation and Alzheimer's research.", 29 "image": [ 30 "https://seaheroquest.com/shq-papers-1-1.png", 31 "https://seaheroquest.com/shq-papers-4-3.png", 32 "https://seaheroquest.com/shq-papers-16-9.png" 33 ] 34 } 35 </script>
35<!----><title>Scientific Papers - Sea Hero Quest</title> 36 <link href="./_app/immutable/assets/0.D6FU2Y3i.css" rel="stylesheet"> 37 <link href="./_app/immutable/assets/PressButton.C25pua9V.css" rel="stylesheet"> 38 <link href="./_app/immutable/assets/Footer.CyxL7Kxa.css" rel="stylesheet"> 39 </head> 40 <body data-sveltekit-preload-data="hover"> 41 <div style="display: contents"><!--[--><!--[0--><!--[--><!--[--><div class="fixed top-0 left-0 right-0 z-50" style="isolation: isolate;"><nav class="mx-auto px-6 py-4 sm:px-6 lg:px-8 items-center transition-all duration-300 bg-dark-blue border-b-2 border-[#2D7798]"><div class="max-w-5xl mx-auto"><div class="flex justify-between items-center"><a href="/" class="flex-shrink-0"><img src="/logo/horizontal.svg" alt="Sea Hero Quest" class="h-[42px]"/></a> <div class="flex items-center gap-[14px]"><a href="https://tally.so/r/3XR5VL" target="_blank" rel="noopener noreferrer" class="hidden md:flex items-center gap-[9px] !rounded-full border-2 border-[#2D7798] hover:border-[#DBF8FF] !px-4 !py-1.5 transition-colors duration-300" style="font-family: 'Lexend', sans-serif;"><img src="/icons/invite.svg" alt="" class="w-[24px] h-[24px] shrink-0"/> <span class="text-[#DBF8FF] text-[18px] font-medium leading-[28px]">Join Waitlist</span></a> <button class="inline-flex items-center justify-center !p-2 !rounded-[10px] border-2 border-[#2D7798] text-[#2D7798] hover:border-[#DBF8FF] hover:text-[#DBF8FF] transition-colors duration-300"><span class="sr-only">Open main menu</span> <div class="flex flex-col justify-center items-center w-6 h-6 gap-[5px]"><span class="block w-5 h-[2px] rounded-full bg-current transition-all duration-300 "></span> <span class="block w-5 h-[2px] rounded-full bg-current transition-all duration-300 "></span> <span class="block w-5 h-[2px] rounded-full bg-current transition-all duration-300 "></span></div></button></div></div></div></nav> <!--[-1--><!--]--></div><!----> <main class="min-h-screen bg-sea-white"><section class="pt-24 pb-16 px-4 sm:px-6 lg:px-8"><div class="max-w-6xl mx-auto"><div class="text-center my-12"><h1 class="text-2xl font-medium leading-tight text-dark-blue m-6 pb-20">Sea Hero Quest in data</h1> <div class="grid grid-cols-1 md:grid-cols-3 gap-8 mb-12"><div class="text-center bg-dark-blue py-12 rounded-panel border-2 border-solid-blue"><div class="text-4xl md:text-5xl font-bold text-primary-major mb-2">0.0m</div> <div class="text-sea-white text-lg">Players built the benchmark</div></div> <div class="text-center bg-dark-blue py-12 rounded-panel border-2 border-solid-blue"><div class="text-4xl md:text-5xl font-bold text-primary-major mb-2">0</div> <div class="text-sea-white text-lg">Research projects running</div></div> <div class="text-center bg-dark-blue py-12 rounded-panel border-2 border-solid-blue"><div class="text-4xl md:text-5xl font-bold text-primary-major mb-2">0</div> <div class="text-sea-white text-lg">Papers published</div></div></div></div></div></section> <section class="py-16 px-4 sm:px-6 lg:px-8"><div class="max-w-6xl mx-auto"><h2 class="text-2xl font-medium leading-tight text-dark-blue mb-8 text-center">Selected highlights</h2> <p class="text-dark-blue/80 text-center mb-12">In no particular order, this is a selection of highlights from published papers, preprints and talks relating to Sea Hero Quest.</p> <div class="space-y-8"><!--[--><div class="bg-white rounded-panel p-6 border-2 border-solid-blue/30"><div class="flex flex-col lg:flex-row gap-6 p-4"><!--[0--><div class="lg:w-1/2 flex-shrink-0"><div class="aspect-video w-full overflow-hidden rounded-panel bg-dark-blue"><iframe class="block h-full w-full" src="https://www.youtube-nocookie.com/embed/uH2z2L7z-Es?rel=0" title="Professor Hugo Spiers on navigation and virtual reality games in Alzheimer's research" allow="accelerometer; autoplay; clipboard-write; encrypted-media; gyroscope; picture-in-picture; web-share" allowfullscreen="" loading="lazy" referrerpolicy="strict-origin-when-cross-origin"></iframe></div></div><!--]--> <div class="flex-1"><h3 class="text-lg font-medium leading-7 text-dark-blue mb-2">Professor Hugo Spiers on navigation and virtual reality games in Alzheimer's research</h3> <p class="text-primary-major font-medium mb-3">UCL Faculty of Brain Sciences, video</p> <p class="text-dark-blue/90 mb-4">Hugo Spiers, one of the scientists behind Sea Hero Quest, explains why the ability to find your way is such a sensitive early signal of Alzheimer's, and how a video game played by millions became a research tool for measuring it.</p> <!--[-1--><!--]--> <!--[-1--><!--]--></div></div></div><div class="bg-white rounded-panel p-6 border-2 border-solid-blue/30"><div class="flex flex-col lg:flex-row gap-6 p-4"><!--[0--><div class="lg:w-1/2 flex-shrink-0"><div class="aspect-video w-full overflow-hidden rounded-panel bg-dark-blue"><iframe class="block h-full w-full" src="https://www.youtube-nocookie.com/embed/wg4PkrDUefs?rel=0&cc_load_policy=1&cc_lang_pref=en" title="Antoine Coutrot, un jeu vidéo pour détecter Alzheimer" allow="accelerometer; autoplay; clipboard-write; encrypted-media; gyroscope; picture-in-picture; web-share" allowfullscreen="" loading="lazy" referrerpolicy="str
41ict-origin-when-cross-origin"></iframe></div></div><!--]--> <div class="flex-1"><h3 class="text-lg font-medium leading-7 text-dark-blue mb-2">Antoine Coutrot, un jeu vidéo pour détecter Alzheimer</h3> <p class="text-primary-major font-medium mb-3">CNRS Talents, video (in French, English captions)</p> <p class="text-dark-blue/90 mb-4">Antoine Coutrot, the CNRS researcher behind much of the Sea Hero Quest data science, on how a video game can help detect Alzheimer's disease early. Part of the CNRS Talents series, in French with English captions turned on by default.</p> <!--[-1--><!--]--> <!--[-1--><!--]--></div></div></div><div class="bg-white rounded-panel p-6 border-2 border-solid-blue/30"><div class="flex flex-col lg:flex-row gap-6 p-4"><!--[1--><div class="lg:w-1/3 flex-shrink-0"><img src="/shq-paper-gameplay.jpg" alt="Research visualization" class="w-full h-auto rounded-panel"/></div><!--]--> <div class="flex-1"><h3 class="text-lg font-medium leading-7 text-dark-blue mb-2">From Players to Participants: Citizen Science and Video Games to Understand Cognition</h3> <p class="text-primary-major font-medium mb-3">Syrine Salouhou, Edgar Dubourg, Maxwell Scott-Slade et al. 2026 (in review)</p> <p class="text-dark-blue/90 mb-4">A timely review on why citizen science video games are becoming essential for cognitive research. The paper lays out the scale of the problem â 96% of psychology participants come from just 12% of the world â and shows how game-based projects like Sea Hero Quest (4M+ players across the globe) help close that gap. It also pulls in rare practical guidance from professional game developers on what it actually takes to build a game that's scientifically rigorous, ethically sound, and engaging enough to keep players coming back.</p> <!--[-1--><!--]--> <!--[0--><a href="https://arxiv.org/abs/2604.24321" target="_blank" rel="noopener noreferrer" class="inline-block px-5 py-2 border-2 border-dark-blue text-dark-blue hover:bg-dark-blue hover:text-sea-white transition-colors duration-200 rounded-pill font-medium">Read Review Paper</a><!--]--></div></div></div><div class="bg-white rounded-panel p-6 border-2 border-solid-blue/30"><div class="flex flex-col lg:flex-row gap-6 p-4"><!--[1--><div class="lg:w-1/3 flex-shrink-0"><img src="/shq-paper-grid-cells.png" alt="Research visualization" class="w-full h-auto rounded-panel"/></div><!--]--> <div class="flex-1"><h3 class="text-lg font-medium leading-7 text-dark-blue mb-2">Spatial navigation in preclinical Alzheimer's disease: A review</h3> <p class="text-primary-major font-medium mb-3">Syrine Salouhou, Victor Gilles, Remi Vallée et al. 2026 (preprint)</p> <p class="text-dark-blue/90 mb-4">This review makes a strong case for what we've believed for years â that spatial navigation is one of the most sensitive early signals of Alzheimer's. The authors map how path integration and wayfinding rely on the exact brain circuits where AD pathology first appears, and show that navigation performance in cognitively unimpaired people correlates with plasma and CSF biomarkers like p-tau. A great synthesis of why scalable navigation testing matters for detecting at-risk individuals before symptoms emerge.</p> <!--[-1--><!--]--> <!--[0--><a href="https://arxiv.org/abs/2603.23082" target="_blank" rel="noopener noreferrer" class="inline-block px-5 py-2 border-2 border-dark-blue text-dark-blue hover:bg-dark-blue hover:text-sea-white transition-colors duration-200 rounded-pill font-medium">Read Review Paper</a><!--]--></div></div></div><div class="bg-white rounded-panel p-6 border-2 border-solid-blue/30"><div class="flex flex-col lg:flex-row gap-6 p-4"><!--[1--><div class="lg:w-1/3 flex-shrink-0"><img src="/shq-paper-2.gif" alt="Research visualization" class="w-full h-auto rounded-panel"/></div><!--]--> <div class="flex-1"><h3 class="text-lg font-medium leading-7 text-dark-blue mb-2">Social determinants of cognitive aging trajectories across 39 countries</h3> <p class="text-primary-major font-medium mb-3">Victor Gilles, Syrine Salouhou, Rémi Vallée et al. 2024 (preprint)</p> <p class="text-dark-blue/90 mb-4">The best descriptions come from the research community â like this short sentence we really liked from an upcoming paper that looks at 'Social determinants of cognitive aging trajectories across 39 countries'.</p> <!--[0--><blockquote class="border-l-4 border-primary-major pl-4 text-dark-blue/80 italic mb-4">
41"Cognition is assessed using Sea Hero Quest, a spatial navigation video game that predicts spatial ability in the real world."</blockquote><!--]--> <!--[0--><a href="https://www.medrxiv.org/content/10.1101/2024.12.05.24318553v1" target="_blank" rel="noopener noreferrer" class="inline-block px-5 py-2 border-2 border-dark-blue text-dark-blue hover:bg-dark-blue hover:text-sea-white transition-colors duration-200 rounded-pill font-medium">Read the paper</a><!--]--></div></div></div><div class="bg-white rounded-panel p-6 border-2 border-solid-blue/30"><div class="flex flex-col lg:flex-row gap-6 p-4"><!--[1--><div class="lg:w-1/3 flex-shrink-0"><img src="/graph.png" alt="Research visualization" class="w-full h-auto rounded-panel"/></div><!--]--> <div class="flex-1"><h3 class="text-lg font-medium leading-7 text-dark-blue mb-2">Geometry of navigation identifies genetic-risk and clinical Alzheimer's disease</h3> <p class="text-primary-major font-medium mb-3">Uzu Lim et al. 2024 (preprint)</p> <p class="text-dark-blue/90 mb-4">We love this pre-print by Uzu Lim because it looks at alternative performance metrics in the existing normative benchmark. In our eyes, it's a really exciting analysis of some additional ways to determine performance from things like path curvature and boundary effects in conjunction with Duration and Length. One to watch!</p> <!--[-1--><!--]--> <!--[0--><a href="https://www.medrxiv.org/content/10.1101/2023.10.01.23296035v1" target="_blank" rel="noopener noreferrer" class="inline-block px-5 py-2 border-2 border-dark-blue text-dark-blue hover:bg-dark-blue hover:text-sea-white transition-colors duration-200 rounded-pill font-medium">Read the paper</a><!--]--></div></div></div><!--]--></div></div></section> <section class="py-16 px-4 sm:px-6 lg:px-8"><div class="max-w-6xl mx-auto"><h2 class="text-2xl font-medium leading-tight text-dark-blue mb-8 text-center">Papers</h2> <div class="bg-white rounded-panel overflow-hidden border-2 border-solid-blue/30"><div class="overflow-x-auto"><table class="w-full"><thead class="bg-dark-blue"><tr><th class="text-left py-4 px-6 text-sea-white font-medium">Year</th><th class="text-left py-4 px-6 text-sea-white font-medium">Paper</th><th class="text-left py-4 px-6 text-sea-white font-medium">Journal</th><th class="text-left py-4 px-6 text-sea-white font-medium">Author(s)</th></tr></thead><tbody class="divide-y-2 divide-solid-blue/15"><!--[--><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2026</td><td class="py-4 px-6"><a href="https://www.sciencedirect.com/science/article/pii/S0028393226001648" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Aphantasia is associated with spatial memory and navigation difficulties in complex virtual environments</a></td><td class="py-4 px-6 text-dark-blue/80">Neuropsychologia</td><td class="py-4 px-6 text-dark-blue/80">Li, A., Coutrot, A., Faromika, T. I. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2026</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s44159-026-00544-3" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Determinants of individual navigation ability</a></td><td class="py-4 px-6 text-dark-blue/80">Nature Reviews Psychology</td><td class="py-4 px-6 text-dark-blue/80">Emre Yavuz & Hugo J. Spiers</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2025</td><td class="py-4 px-6"><a href="https://www.cell.com/iscience/fulltext/S2589-0042(25)01702-X" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Separable cognitive and motor decline tracked across adult life-span for goal-directed navigation.</a></td><td class="py-4 px-6 text-dark-blue/80">iScience</td><td class="py-4 px-6 text-dark-blue/80">Gian Luca Lancia, Marco D'Alessandro, Mattia Eluchans. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2025</td><td class="py-4 px-6"><a href="https://direct.mit.edu/opmi/article/doi/10.1162/opmi.a.13/132235/Education-Is-Positively-and-Causally-Linked-With" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Education Is Positively and Causally Linked With Spatial Navigation Ability Across the Lifespan</a></td><td class="py-4 px-6 text-dark-blue/80">Open Mind</td><td class="py-4 px-6 text-dark-blue/80">Antoine Coutrot, Rogier A Kievit, Stuart J Ritchie et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41598-024-52662-8" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Shorter self-reported sleep duration is associated with worse virtual spatial navigation performance in men.</a></td><td class="py-4 px-6 text-dark-blue/80">Scientific Reports</td><td class="py-4 px-6 text-dark-blue/80">Yavuz, E., Gahnstrom, C.J., Goodroe, S. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://www.mdpi.com/2076-3425/14/9/898" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Spatial Orientation Assessment in the Elderly: A Comprehensive Review of Current Tests</a></td><td class="py-4 px-6 text-dark-blue/80">Brain Sciences</td><td class="py-4 px-6 text-dark-blue/80">Panagiota Tragantzopoulou & Vaitsa Giannouli</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://onlinelibrary.wiley.com/doi/10.1002/hbm.26623" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Lost in space(s): Multimodal neuroimaging of disorientation along the Alzheimer's disease continuum</a></td><td class="py-4 px-6 text-dark-blue/80">Human Brain Mapping</td><td class="py-4 px-6 text-dark-blue/80">Peters-Founshtein, G., Gazit, L., Naveh, T. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://www.tandfonline.com/doi/full/10.1080/13875868.2024.2319735" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Individuals learning to drive solo before age 18 have superior spatial navigation ability compared with those who learn later</a></td><td class="py-4 px-6 text-dark-blue/80">Spatial Cognition & Computation</td><td class="py-4 px-6 text-dark-blue/80">Yavuz, E., Manley, E., Gahnstrom, C. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://www.sciencedirect.com/science/article/pii/S0272494424000690?via%3Dihub" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Video gaming, but not reliance on GPS, is associated with spatial navigation performance</a></td><td class="py-4 px-6 text-dark-blue/80">Journal of Environmental Psychology</td><td class="py-4 px-6 text-dark-blue/80">Yavuz, E., Chuanxiuyue He, Gahnstrom, C. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://www.cell.com/iscience/fulltext/S2589-0042%2824%2901199-4?%5FreturnURL=https://linkinghub.elsevier.com/retrieve/pii/S2589004224011994?showall%3Dtrue" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Combining patient-lesion and big data approaches to reveal hippocampal contributions to spatial memory and navigation.</a></td><td class="py-4 px-6 text-dark-blue/80">iScience</td><td class="py-4 px-6 text-dark-blue/80">Pishdadian, S., Coutrot, A., Webber, L. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41562-024-01878-9" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Using games to understand the mind</a></td><td class="py-4 px-6 text-dark-blue/80">Nature Human Behaviour</td><td class="py-4 px-6 text-dark-blue/80">Allen, K., Brändle, F., Botvinick, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2024</td><td class="py-4 px-6"><a href="https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0298116" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">The relationship between object-based spatial ability and virtual navigation performance</a></td><td class="py-4 px-6 text-dark-blue/80">PLoS ONE</td><td class="py-4 px-6 text-dark-blue/80">Garg, T., Velasco, P. F., Patai, Z. E. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2023</td><td class="py-4 px-6"><a href="https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0282255" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Wayfinding and path integration deficits detected using a virtual reality mobile app in patients with traumatic brain injury</a></td><td class="py-4 px-6 text-dark-blue/80">PLoS ONE</td><td class="py-4 px-6 text-dark-blue/80">Seton, C., Coutrot, A., Hornberger, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2023</td><td class="py-4 px-6"><a href="https://doi.org/10.1016/j.cortex.2023.01.001" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Hydrocephalus: A neuropsychological and theoretical primer</a></td><td class="py-4 px-6 text-dark-blue/80">Cortex</td><td class="py-4 px-6 text-dark-blue/80">Zaksaite, T., Loveday, C., Edginton, T. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2023</td><td class="py-4 px-6"><a href="https://researchportal.northumbria.ac.uk/en/publications/motivational-factors-for-participating-in-citizen-science-games" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Motivational Factors for Participating in Citizen Science Games</a></td><td class="py-4 px-6 text-dark-blue/80">Northumbria Research Portal</td><td class="py-4 px-6 text-dark-blue/80">Dalton, R., Wiener, J., Yesiltepe, D. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2023</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41598-023-30937-w" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Cultural determinants of the gap between self-estimated navigation ability and wayfinding performance: evidence from 46 countries</a></td><td class="py-4 px-6 text-dark-blue/80">Scientific Reports</td><td class="py-4 px-6 text-dark-blue/80">Walkowiak, S., Coutrot, A., Hegarty, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2023</td><td class="py-4 px-6"><a href="https://direct.mit.edu/jocn/article/35/3/452/114450/Landmark-dependent-Navigation-Strategy-Declines" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Landmark-dependent Navigation Strategy Declines across the Human Life-Span: Evidence from Over 37,000 Participants</a></td><td class="py-4 px-6 text-dark-blue/80">Journal of Cognitive Neuroscience</td><td class="py-4 px-6 text-dark-blue/80">West, G., Patai, Z. E., Coutrot, A. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2022</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41467-022-34624-8" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Reported sleep duration reveals segmentation of the adult life-course into three phases</a></td><td class="py-4 px-6 text-dark-blue/80">Nature Communications</td><td class="py-4 px-6 text-dark-blue/80">Coutrot, A., Lazar, A. S., Richards, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2022</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41598-022-17634-w" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Predicting real world spatial disorientation in Alzheimer's disease patients using virtual reality navigation tests</a></td><td class="py-4 px-6 text-dark-blue/80">Scientific Reports</td><td class="py-4 px-6 text-dark-blue/80">Puthusseryppady, V., Morrissey, S., Spiers, H. J. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2022</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41398-022-02237-w" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Developments in scalable strategies for detecting early markers of cognitive decline</a></td><td class="py-4 px-6 text-dark-blue/80">Translational Psychiatry</td><td class="py-4 px-6 text-dark-blue/80">Whelan, R., Barbey, F. M., Cominetti, M. R. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2022</td><td class="py-4 px-6"><a href="https://www.nature.com/articles/s41586-022-04486-7" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Entropy of city street networks linked to future spatial navigation ability</a></td><td class="py-4 px-6 text-dark-blue/80">Nature</td><td class="py-4 px-6 text-dark-blue/80">Coutrot, A., Manley, E., Goodroe, S. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2022</td><td class="py-4 px-6"><a href="https://discovery.ucl.ac.uk/id/eprint/10150148/" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Exosomatic Route Choice in Navigation: Evidence from video game player data</a></td><td class="py-4 px-6 text-dark-blue/80">Proceedings of the 13th Space Syntax Symposium</td><td class="py-4 px-6 text-dark-blue/80">McElhinney, S., Zisch, F., Hornsberger, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2022</td><td class="py-4 px-6"><a href="https://pmc.ncbi.nlm.nih.gov/articles/PMC9178368/" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">How the environment shapes our ability to navigate</a></td><td class="py-4 px-6 text-dark-blue/80">Clinical and Translational Medicine</td><td class="py-4 px-6 text-dark-blue/80">Spiers, H. J., Coutrot, A., & Hornberger, M.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2021</td><td class="py-4 px-6"><a href="https://onlinelibrary.wiley.com/doi/full/10.1111/tops.12590" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Explaining World-Wide Variation in Navigation Ability from Millions of People: Citizen Science Project Sea Hero Quest</a></td><td class="py-4 px-6 text-dark-blue/80">
41Topics in Cognitive Science</td><td class="py-4 px-6 text-dark-blue/80">Spiers, H. J., Coutrot, A., & Hornberger, M.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2021</td><td class="py-4 px-6"><a href="https://ieeexplore.ieee.org/abstract/document/9287403" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Capturing and explaining trajectory singularities using composite signal neural networks</a></td><td class="py-4 px-6 text-dark-blue/80">28th European Signal Processing Conference</td><td class="py-4 px-6 text-dark-blue/80">Dubois, H., Le Callet, P., Hornberger, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2021</td><td class="py-4 px-6"><a href="https://www.tandfonline.com/doi/full/10.1080/13875868.2020.1830993" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Computer models of saliency alone fail to predict subjective visual attention to landmarks during observed navigation</a></td><td class="py-4 px-6 text-dark-blue/80">Spatial Cognition & Computation</td><td class="py-4 px-6 text-dark-blue/80">Yesiltepe, D., Ozbil Torun, A., Coutrot, A. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2021</td><td class="py-4 px-6"><a href="https://academic.oup.com/braincomms/article/3/2/fcab087/6248086" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Memory precision of object-location binding is unimpaired in APOE ε4-carriers with spatial navigation deficits</a></td><td class="py-4 px-6 text-dark-blue/80">Brain Communications</td><td class="py-4 px-6 text-dark-blue/80">Gellerson, H. M., Coughlan, G., Hornberger, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2020</td><td class="py-4 px-6"><a href="https://alz-journals.onlinelibrary.wiley.com/doi/abs/10.1002/alz.043085" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Navigation ability before and after menopause</a></td><td class="py-4 px-6 text-dark-blue/80">Alzheimer's & Dementia: The Journal of the Alzheimer's association</td><td class="py-4 px-6 text-dark-blue/80">Chapin, B.A., Khondoker, M., Coughlan, G. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2020</td><td class="py-4 px-6"><a href="https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0239077" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Test-retest reliability of spatial navigation in adults at-risk of Alzheimer's disease</a></td><td class="py-4 px-6 text-dark-blue/80">PLoS ONE</td><td class="py-4 px-6 text-dark-blue/80">Coughlan, G., Puthusseryppady, V., Lowry, E. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2019</td><td class="py-4 px-6"><a href="https://www.pnas.org/doi/abs/10.1073/pnas.1901600116" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Toward personalized cognitive diagnostics of at-genetic-risk Alzheimer's disease</a></td><td class="py-4 px-6 text-dark-blue/80">PNAS</td><td class="py-4 px-6 text-dark-blue/80">Coughlan, G., Coutrot, A., Khondoker, M. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2019</td><td class="py-4 px-6"><a href="https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0213272" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Virtual navigation tested on a mobile app is predictive of real-world wayfinding navigation performance</a></td><td class="py-4 px-6 text-dark-blue/80">PLoS ONE</td><td class="py-4 px-6 text-dark-blue/80">Coutrot, A., Schmidt, S., Coutrot, L. et al.</td></tr><tr class="hover:bg-sea-white/40 transition-colors"><td class="py-4 px-6 text-dark-blue font-medium">2018</td><td class="py-4 px-6"><a href="https://www.sciencedirect.com/science/article/pii/S0960982218307711?via%3Dihub" target="_blank" rel="noopener noreferrer" class="text-primary-major hover:underline font-medium">Global determinants of navigation ability</a></td><td class="py-4 px-6 text-dark-blue/80">Current Biology</td><td class="py-4 px-6 text-dark-blue/80">Coutrot, A., Silva, R., Manley, E. et al.</td></tr><!--]--></tbody></table></div></div></div></section></main> <footer class="bg-dark-blue text-white py-12 border-t-2 border-solid-blue"><div class="max-w-5xl mx-auto px-4 sm:px-6 lg:px-8">
41<div class="grid md:grid-cols-5 gap-8"><div class="col-span-2"><div class="flex items-center gap-2 mb-4"><div class="w-8 h-8 border-2 border-white rounded-lg flex items-center justify-center"><img src="/shq-anchor.svg" alt="Sea Hero Quest" class="w-5 h-5"/></div> <img src="/logo/horizontal.svg" alt="Sea Hero Quest" class="h-[28px]"/></div> <div class="text-sm text-white/30"><p class="mb-2">'Sea Hero Quest' and 'GLITCHERS' are registered trademarks of <a href="https://glitchers.com/" target="_blank" rel="noopener noreferrer" class="hover:text-white/50">GLITCHERS LTD</a>.</p> <p>© 2026 <a href="https://glitchers.com/" target="_blank" rel="noopener noreferrer" class="hover:text-white/50">GLITCHERS LTD</a></p> <p class="mt-2 text-xs text-white/20">v1.9.0 · build 13 (1330f6f)</p></div></div> <div><h4 class="font-semibold mb-4">Research</h4> <ul class="space-y-2 text-white/50"><li><a href="/papers" class="hover:text-white">Read the Papers</a></li> <li><a href="https://celest.seaheroquest.com/" class="hover:text-white">Researcher Portal</a></li></ul></div> <div><h4 class="font-semibold mb-4">Media</h4> <ul class="space-y-2 text-white/50"><li>
41<a href="https://www.figma.com/buzz/SN0rkHzALCUubKPHxlzfrK/SHQ-Assets?node-id=23-2&t=mTgPJnKrUx1Rt9oc-1" class="hover:text-white">Media Pack</a></li> <li><a href="https://glitchers.com/contact" class="hover:text-white">Contact</a></li></ul></div> <div><h4 class="font-semibold mb-4">Legal</h4> <ul class="space-y-2 text-white/50"><li><a href="/privacy-policy" class="hover:text-white">Privacy Policy</a></li> <li><a href="/terms" class="hover:text-white">Terms of Service</a></li> <li><a href="/ethics" class="hover:text-white">Ethics</a></li></ul></div></div></div></footer><!----><!--]--><!----><!--]--><!--]--> <!--[-1--><!--]--><!--]--> 42 43
43<script> 44 { 45 __sveltekit_11eiagi = { 46 base: new URL(".", location).pathname.slice(0, -1) 47 }; 48 49 const element = document.currentScript.parentElement; 50 51 Promise.all([ 52 import("./_app/immutable/entry/start.BQUWCIEl.js"), 53 import("./_app/immutable/entry/app.CHxdJccT.js") 54 ]).then(([kit, app]) => { 55 kit.start(app, element, { 56 node_ids: [0, 18], 57 data: [null,null], 58 form: null, 59 error: null 60 }); 61 }); 62 } 63 </script>
63 64 </div> 65
65<script type="module" src="https://static.cloudflareinsights.com/beacon.min.js/v31edd6df95cf4e85bb4c19e7a9bdbcba1788362987495" integrity="sha512-iIg7k2xntmwu6/uSb5tpc/hySgZc4eoL31yB29W6tJFo2akwjPWcEqnCEdJvGexCL0KEQwVYv5BlowfhVz26hg==" data-cf-beacon='{"version":"2024.11.0","token":"c41824a5175843efb99a06fdf15a7017","r":1,"spa":2}' crossorigin="anonymous"></script>
65 66</body> 67</html>
Line numbers count LF bytes from the start of the resource, as the search results do. Vendor segments are library code the classifier recognised; they are stored but not indexed. Bytes are shown as Latin1 characters, one per byte.